SNP leader Nicola Sturgeon was the second leader to take to the stage.

As she wants a confirmatory vote on Brexit, she was asked whether that would also apply to any Scottish independence deal. You can watch that clip below.

Image source, EPA

Jeremy Corbyn

• The Labour leader was up first to take questions. He said he would "remain neutral" in the Brexit referendum planned by Labour and not campaign for Leave or Remain.
• He was also challenged on anti-Semitism faced by Labour MPs, including Ruth Smeeth, who is Jewish.

Nicola Sturgeon

• The SNP leader and Scottish first minister was asked about her red lines in the event of another hung Parliament, saying she would not work with the Tories.
• But she said she was prepared to make a deal with Labour if Jeremy Corbyn committed to end austerity and agreed to hold another Scottish independence referendum.

Jo Swinson

• The Liberal Democrat leader was asked whether she regretted saying she could become prime minister - given that her party only had 20 MPs at the end of the last Parliament.
• She was also grilled on austerity and her record in government while in coalition with the Conservatives between 2010 and 2015.

Boris Johnson

• The prime minister was challenged on comments he made in articles as a journalist - the now infamous "letter boxes" article he wrote was highlighted by presenter Fiona Bruce. Mr Johnson defended his "right to speak out".
• He was also pushed on issues of trust, including a question on promises not fully realised by the Tories to recruit more GPs. Pressed on the release of the Russian interference report, Mr Johnson said he knew of "no evidence" of interference in any UK electoral event.

Over in the spin room, we are hearing reaction to the leaders' performances in the debate.

On the side of Boris Johnson is Culture Secretary Nicky Morgan, who is not standing in this election. She's asked about the issue of trust being put repeatedly to Mr Johnson, but almost immediately turns the question around to Brexit.

Ms Morgan says Brexit is the big issue of the day and the PM offers a much clearer position than Jeremy Corbyn.

"That’s the contrast between the two men who are actually vying to be prime minister," she says.

She describes Jeremy Corbyn's comment that he would "remain neutral" in the Brexit referendum planned by Labour and not campaign for Leave or Remain as "extraordinary".

"Why would the EU give new terms for a new deal if the new leader was not going to advocate for that deal?" she asks.

"So on the biggest issue of the day Jeremy Corbyn does not have a position. A big contrast to Boris Johnson’s position, which has been very clear."

The plight of the "Waspi women" was raised by the audience as the quizzed Boris Johnson, but who are they? Waspi stands for "Women Against State Pension Inequality" and they are a group who are campaigning against changes to women's state pension age.

The retirement age for women rose from 60 to 65, in line with men, and will go up to 66 by 2020, and to 67 by 2028.

Women born in the 1950s claim the rise is unfair because they were not given enough time to make adjustments to cope with years without a state pension.

In total around 2.6 million women were affected by the 2011 changes. Read more about the background to the campaign here.

Iain Watson
Political correspondent

These were well-informed questions from an educated audience. Intelligent hostility, shall we say.

Jeremy Corbyn did fairly well - but he gave away a bit more on the issue of Scottish independence, and he also said out loud that he'd be neutral in another Brexit referendum. The Tories may use that.

Nicola Sturgeon talked more about how she would try to exert leverage against Mr Corbyn.

Jo Swinson got a pretty hard time, over Brexit and her record in government. Boris Johnson found himself unable to keep coming back to Brexit and was pushed on a whole host of other issues.

Boris Johnson says he "over-delivered" in his time as London mayor - and is already delivering on the promises he made when entering No 10 - recruiting 20,000 more police officers, for example.

He says he understands the "massive pressures" the NHS faces and is already putting more money in.

Mr Johnson brings it back to Brexit again - and again draws some jeers for doing so - because he says the economy will continue to be strangled until it is done.

There's then some clapping as he says people still haven't got what they voted for - he says he has "an oven-ready deal", waiting to go but all the other parties are "trying to frustrate" the process.

The Tory leader then leaves the stage to some cheers.

Our general election policy guide is a concise summary of where the parties stand on big issues like Brexit, education and the NHS – and you can easily compare what each party is promising to do if they get elected.

The next question is on promises made by the Conservatives to recruit more GPs - promises not fully realised - so how, the questioner asks, can we trust you?

Mr Johnson says the number of GPs has gone up - "but of course, I want more GPs, I want more investment in the NHS," he adds.

There's then some wrangling over the number of hospitals the Tories say they're going to build.

Boris Johnson seems to smile slightly as he knows this is a contentious issues, but he insists that ultimately the 40 new hospitals he keeps referring to will be built.

Another questioner now talks about how much strain NHS workers, himself included, are under.

Boris Johnson says the audience member is "right", and he is trying to recruit more staff.

"The only way we can do that is to have a strong economy..." he says, and tells the audience he doesn't believe they would get that under Jeremy Corbyn.

Iain Watson
Political correspondent

It is interesting that so far, Boris Johnson has not been allowed to drift back to Brexit when in hotter water and is being challenged on the scale of his promised hospital programme and the NHS more widely.

Reality Check

Boris Johnson said he would raise the starting threshold of national insurance to £9,500 in 2020-21, saving people £100 per year.

The Institute for Fiscal Studies (IFS) says the saving would be slightly less, £85 per year, external (£1.63 a week).

Boris Johnson says his party has "zero tolerance of Islamophobia" and will have an inquiry into all forms of prejudice in the Conservative Party.

He says he is "proud" of his party's record on promoting women - "which party has produced two female prime ministers?" he asks.

"Of course there is more we can do," he goes on, but says the gender pay gap has shrunk under the Tories and he is making announcements on childcare too.

Iain Watson
Political correspondent

Boris Johnson challenged on racist rhetoric - there is derisive laughter when he says you can find things which may look offensive.

There is no applause when he claims he was really standing up for freedom of women to wear what they want.

Boris Johnson is asked whether he has personally contributed to discrimination and prejudice against minorities.

Fiona Bruce lists some examples, including the now infamous "letter boxes" article he wrote, in which he said full-face veils should not be banned, but it was “absolutely ridiculous” women chose to “go around looking like letterboxes”.

"I've genuinely never intended to cause hurt or pain to anybody," the Conservative leader replies.

He says he defends his "right to speak out", although he accepts that some of his words could appear "offensive" if taken out of context.

Mr Johnson says the article was actually making "a strong liberal defence of a woman's right to wear whatever she chooses".

He says he has always campaigned on gender equality - including while as foreign secretary when he led efforts to improve education for girls around the world.

Iain Watson
Political correspondent

Boris Johnson on a stickier wicket alleging no Russian interference in politics while not promising to publish the report on it pre-election.

He is very keen to get back to Brexit despite barracking by one audience member in particular.

Why should a young person vote Conservative? a student in the audience asks.

We are the party who are going to help you get a highly paid job, we're going to help you get on the housing ladder, Boris Johnson says.

His claims about the number of houses the Tories have been building are met with some sceptical jeers, but Mr Johnson presses on and defends his party's record.

He's then pushed hard on why he won't release a report into allegations of Russian interference in UK democracy. There's clearly strong feeling in the audience about this.

"There is absolutely no evidence as far as I'm aware..." Boris Johnson starts to say.

He says he can "see no reason to interfere with the normal timetable" for the release of the report.

The questioner is angry at that - "it's not true", he says, pressing for more.

"This is complete Bermuda Triangle stuff," Boris Johnson concludes.

Earlier we heard Jeremy Corbyn tell the Question Time audience that if he becomes prime minister he will remain neutral in the campaign for a further referendum on Brexit.

You can watch the clip below, or read our full story here.

A questioner asks about her own struggles to make ends meet and what reassurances she can get from Boris Johnson that things won't get worse.

"Of course we want to deal with the expression of poverty but also the causes of poverty," he replies, and says that's why he wants to invest in education, child services, and put more money into people's pockets to help with the cost of living.

That's the idea behind his National Insurance cut - announced earlier this week.

Mr Johnson acknowledges it's not a lot of money initially but adds: "We want to do more."

He says he believes Universal Credit has worked to help hundreds of thousands of people back into work and that is the aspect of it he wants to focus on.

Iain Watson
Political correspondent

The Conservatives' own polling suggests that apart from Brexit, their target voters want a bit of 'sunlit uplands', so he peppers his address with 'ambitions' for the country.

Boris Johnson is asked about rising food bank use and school funding shortages under the Conservatives.

He says he talked to many doctors, teachers, and other public servants since becoming PM and he is trying to "level up" funding and put more money into schools.

He says he raised the living wage while London mayor and then George Osborne "nicked" the idea and made it a national policy.

He insists he does want to tackle poverty and "unleash the potential" of every child.

Iain Watson
Political correspondent

Well, as they say - tough crowd. I have always said audience questions can be more challenging than conventional debates.

Mind you, Boris Johnson has the benefit of going last and is able to attack what Jeremy Corbyn said earlier.

And the PM's delivery has passion and avoids insouciance.