Sam Warburton and his men are lining up in the bowels of the Millennium Stadium, dragon-like flamers are flaring up around the pitch, but here's who will be entertaining us this afternoon:

Wales: Halfpenny, Cuthbert, J. Davies, Roberts, L. Williams, Biggar, Webb, Jenkins, Baldwin, Lee, Ball, A. Jones, Lydiate, Warburton, Faletau.

Replacements: E. Phillips, Jarvis, R. Jones, Charteris, King, M. Phillips, Priestland, S. Williams.

South Africa: le Roux, Hendricks, Serfontein, de Villiers, Mvovo, Lambie, Reinach, Mtawarira, B. du Plessis, Oosthuizen, Etzebeth, Matfield, Coetzee, Mohoje, Vermeulen.

Replacements: Strauss, Nyakane, Redelinghuys, de Jager, Carr, Hougaard, Pollard, de Allende.

Referee: John Lacey (Ireland)

England field yet another re-jigged centre partnership as Billy Twelvetrees is recalled to play alongside Brad Barritt.

Dylan Hartley and Tom Wood come back into the pack but the biggest call is probably at fly-half.

George Ford starts at 10 for the second game in a row with Owen Farrell, who was moved to 12 last weekend, relegated to the bench.

Wales coach Warren Gatland, speaking on BBC One: "We've had a good look at South Africa. I'm not sure the best team turned up against Ireland in Dublin, and they got turned over. We have to make sure our kicking game's good.

"We've spoken this week and the theme has been that we have to keep going every moment for 80 minutes. I can't fault the effort last week but one or two individuals switched off for a moment or two, and we have to make sure we don't do that."

Northampton wing George North, ruled out of Wales' match against South Africa after suffering a blow to the head against New Zealand last Saturday:, external Gutted not to be playing today! But best of luck to the Welsh Rugby Union men! Wales by 7!! Come on boys!

England: Brown; Watson, Barritt, Twelvetrees, May; Ford, Youngs; Marler, Hartley, Wilson, Attwood, Lawes, Wood, Robshaw, Morgan.

Replacements: Webber, Mullan, Brookes, Kruis, Haskell, Wigglesworth, Farrell, Yarde.

Australia: Folau; Speight, Ashley-Cooper, Toomua, Horne; Foley, Phipps; Slipper, Faingaa, Kepu, Carter, Simmons, McMahon, Hooper, McCalman.

Replacements: Hanson, Robinson, Alexander, Skelton, Jones, White, Cooper, Beale.

Referee: Jerome Garces (France)

Martyn Williams
Former Wales captain on BBC One

"Wales have been answering the same questions, week in, week out, for the last five or six years: Why do we let things slip in the last 10 minutes? I'm hoping you'll see a desperate, frustrated Wales team today putting some questions to bed. But they'll need a performance against a very big South Africa side."

South Africa coach Heyneke Meyer has gone in front of the cameras and predicts: "it's going to be a real test of character for our team".

"I truly believe they've been unlucky," he says of Wales' record against the southern hemisphere teams. Even in South Africa they've been unlucky. They're a great side, but we have to control what we can control and take it to them and be physical."

Image source, @EnglandRugby

England Rugby:, external Bats have been placed by the England and Australia benches as marks of respect for Phillip Hughes. #putoutyourbats

Australia Test cricketer Hughes, 25, died on Thursday after he suffered a blow to the neck during a domestic match in Sydney.

Ian Robertson
BBC rugby union correspondent on Radio 5 live sports extra

"I don't think England know what their best centre pairing is. If you were to ask six ex-internationals for their selections, you would probably get five different pairings.

"At Bath, George Ford pairs well with Kyle Eastmond and Jonathan Joseph. With England, I think Stuart Lancaster will always put down Mani Tuilagi; he's strong, he has a low centre of gravity. One of those in every team is great. Luther Burrell would probably be the one Stuart would like in there, but he's been injured, so he's given Billy Twelvetrees a last chance. He's not really fulfilled his potential at international level."

Lewis Moody
Former England captain on BBC Radio 5 live sports extra

"England have to win today. Over the last three weeks, they have left too many points on the pitch. George Ford is an exciting selection. I don't think there's a lack of attacking talent in this side, but how are they going to get the ball to them?"

Huge - that's how Lewis Moody describes it and it's hard to disagree. England need to win today to reignite belief they are on an upward trajectory.

These two sides meet in the World Cup in less than a year's time.

Today's game is not relevant, say the players and coaches. The fans? They're not to sure - there's surely a big psychological marker to be laid down today.

Owen Farrell is out of the picture, for now at least, but who is the best home nations fly-half? George Ford showed promise against Samoa, while Dan Biggar has a controlling influence over Wales - but there's also Jonny Sexton and Scotland's Finn Russell to think of. So, who would be your pick and why?

Tweet us using the hashtag #bbcrugby or text 81111 - with your name and RUGBY before your message. Alternatively, you could write your thoughts on BBC Sport's Facebook page.

Image source, PA

There's no Owen Farrell in the England line-up this afternoon with the Saracens man making way for Billy Twelvetrees. How did Farrell react to being overlooked for the first time since June 2012? "His reaction is very much what you would expect from a professional rugby player... not accepting it is the right thing," said backs coach Andy Farrell of his son's demotion.

There's too much going on for one keyboard so James Standley has warmed up, slapped some Vaseline over his eyebrows, gone through what looked like of war dance and is ready to join the action, guiding you through all that's going on at Twickenham.

The Guardian's , externalPaul Rees writes of Wales' late, late defeats against the big three: "It is at the point when a match reaches the stage when fatigue, mental and physical, sets in, and it becomes about what lies deep within, that Wales lose their grip. They are vulnerable when a Test swivels on playing rather than coaching."

Image source, PA

"You cannot overemphasise how important this game against Australia is. It is huge," says Lewis Moody in his column in the Independent. , external"The importance of ending the autumn campaign on a high and going into the Six Nations remembering a win, and playing with confidence, really cannot be overstated."

Other opinions and other media outlets are available.

Paul Grayson
Former England fly-half on BBC Radio 5 live sports extra

"It's the last of the four Autumn internationals for England, and by now, you should be seeing something cohesive. You need to be at something like your best, and England have been nowhere near it so far."

England's record against Southern Hemisphere teams isn't as pitiful as their neighbours Wales, and Stuart Lancaster's men might be confident of victory against a team which has lost five of their last six away games.

"We have spoken about the pressure that can build, especially with a home World Cup," captain Chris Robshaw said this week. "And we definitely feel that all the outside stuff has built up more this autumn than in previous campaigns.

"But it is about embracing that. We have got a fantastic opportunity to go out there on Saturday and do something special."

Image source, Getty Images

Alastair Eykyn
BBC Radio 5 live sports extra commentator at Twickenham

"Today, almost for the first time since he took over, Stuart Lancaster is as tight as a drum. He looks nervous. This is a huge game for him."

Bad news for those still making their way to HQ because there's major travel disruption for many fans travelling south on the railways to Twickenham after a police incident at Watford Junction., external

BBC Radio 5 live Breakfast presenter Rachel Burden,, external who is trying to get to the game, has been live tweeting:

"Not great news for Eng fans heading for Euston. We're all stuck due to incident further up the line. Going to be very tight to get there.

"Moving very slowly now. Any advice on the quickest route from Euston to Twickenham? Had planned tube and train...

"We've made friends with fellow travellers so it's all for one and one for all now. Handy for shared taxi fares!"

Image source, PA